Preface to the Princeton Landmarks in Biology Edition vii Preface xi Symbols Used xiii 1. The Importance of Islands 3 2. Area and Number of Speicies 8 3. Further Explanations of the Area-Diversity Pattern 19 4. The Strategy of Colonization 68 5. Invasibility and the Variable Niche 94 6. Stepping Stones and Biotic Exchange 123 7. Evolutionary Changes Following Colonization 145 8. Prospect 181 Glossary 185 References 193 Index 201

The Theory of Island Biogeography

The cultural origins of human cognition

The revolution that wasn't: a new interpretation of the origin of modern human behavior.

Bones of recent mammals in the Amboseli Basin, southern Kenya, exhibit distinctive weathering characteristics that can be related to the time since death and to the local conditions of temperature, humidity and soil chemistry. A categorization of weathering characteristics into six stages, recognizable on descriptive criteria, provides a basis for investigation of weathering rates and processes. The time necessary to achieve each successive weathering stage has been calibrated using known-age carcasses. Most bones decompose beyond recognition in 10 to 15 yr. Bones of animals under 100 kg and juveniles appear to weather more rapidly than bones of large animals or adults. Small-scale rather than widespread environmental factors seem to have greatest influence on weathering characteristics and rates. Bone weathering is potentially valuable as evidence for the period of time represented in recent or fossil bone assemblages, in- cluding those on archeological sites, and may also be an important tool in censusing populations of animals in modern ecosystems.

Taphonomic and Ecologic Information Form Bone Weathering

Wonderful Life: The Burgess Shale and the Nature of History, Stephen Jay Gould. W. W. Norton, New York (1989), 347, Price $19.95 (U.S.A.), $27.95 (Canada)

http://www.u.arizona.edu/~mstiner/pdf/Stiner_etal1995.pdf

Differential Burning, Recrystallization, and Fragmentation of Archaeological Bone

Human beings depend more on technology than any other animal-the use of tools and weapons is vital to the survival of our species. What processes of biocultural evolution led to this unique dependence? Steven Kuhn turns to the Middle Paleolithic (Mousterian) and to artifacts associated with Neandertals, the most recent human predecessors. His study examines the ecological, economic, and strategic factors that shaped the behaviour of Mousterian tool makers, revealing how these hominids brought technological knowledge to bear on the basic problems of survival. Kuhn's main database consists of assemblages of stone artifacts from four caves and a series of open-air localities situated on the western coast of the Italian peninsula. Variations in the ways stone tools were produced, maintained, and discarded demonstrate how Mousterian hominids coped with the problems of keeping mobile groups supplied with the artifacts and raw materials they used on a daily basis. Changes through time in lithic technology were closely tied to shifting strategies for hunting and collecting food. Some of the most provocative findings of this study stem from observations about the behavioural flexibility of Mousterian populations and the role of planning in foraging and technology.

Mousterian Lithic Technology: An Ecological Perspective

It is widely believed that the design of transported artifacts and toolkits employed by mobile populations is influenced by two main factors, portability and potential utility. Choices about the kinds of artifacts to carry around can be modeled as an optimization problem, in which it is beneficial to obtain the greatest potential utility for the minimum total weight. This study approaches the problem analytically, making a few simple assumptions about artifact geometry and the relations between utility and artifact size. If artifact utility is calculated as a function of potential for renewal, transported toolkits should consist entirely of relatively small finished tools. Moreover, most gains in durability or multifunctionality that require increases in overall size are outweighed by increased transport cost. Inconsistencies between these expectations and ethnographic and archaeological observations point to situations in which artifact functionality is more closely constrained by overall size or mass.

A Formal Approach to the Design and Assembly of Mobile Toolkits

Despite the rapid expansion of archaeological knowledge of the Paleolithic over the past several decades, some generalized interpretive frameworks inherited from previous generations of researchers are remarkably tenacious. One of the most persistent of these is the assumed correlation between blade technologies, Upper Paleolithic industries, and anatomically (and behaviorally) modern humans. In this paper, we review some of the evidence for the production of early blade technologies in Eurasia and Africa dating to the late Lower and the Middle Paleolithic. The basic techniques for blade production appeared thousands of years before the Upper Paleolithic, and there is no justification for linking blades per se to any particular aspect of hominid anatomy or to any major change in the behavioral capacities of hominids. It is true that blades came to dominate the archaeological records of western Eurasia and Africa after 40,000 years ago, perhaps as a consequence of increasing reliance on complex composite tools during the Upper Paleolithic. At the same time, evidence from other regions of the world demonstrates that evolutionary trends in Pleistocene Eurasia were historically contingent and not universal. [Middle Paleolithic, Upper Paleolithic, blade technology, human evolution, hominid behavior and capacities]

The Big Deal about Blades: Laminar Technologies and Human Evolution

Recent huntergatherers display much uniformity in the division of labor along the lines of gender and age. The complementary economic roles for men and women typical of ethnographically documented huntergatherers did not appear in Eurasia until the beginning of the Upper Paleolithic. The rich archaeological record of Middle Paleolithic cultures in Eurasia suggests that earlier hominins pursued more narrowly focused economies, with womens activities more closely aligned with those of men with respect to schedule and ranging patterns than in recent forager systems. More broadly based economies emerged first in the early Upper Paleolithic in the eastern Mediterranean region and later in the rest of Eurasia. The behavioral changes associated with the Upper Paleolithic record signal a wider range of economic and technological roles in forager societies, and these changes may have provided the expanding populations of Homo sapiens with a demographic advantage over other hominins in Eurasia.

http://www.u.arizona.edu/~mstiner/pdf/Kuhn_Stiner2006.pdf

Steven L. Kuhn

Papers

Differential Burning, Recrystallization, and Fragmentation of Archaeological Bone

Mousterian Lithic Technology: An Ecological Perspective

A Formal Approach to the Design and Assembly of Mobile Toolkits

The Big Deal about Blades: Laminar Technologies and Human Evolution

What's a Mother to Do? The Division of Labor among Neandertals and Modern Humans in Eurasia